Date: 2008-10-26 20:39:03

Alejandro Estrada

Guatemalan black howler

Mantled howler

Venezuelan red howler

Black howler

Spider monkey

Primate

Monkey

Atelidae

Howler monkeys

Zoology

Fauna of South America

American Journal of Primatology 70:1–[removed]RESEARCH ARTICLE Group Size and Composition Influence Male and Female Reproductive Success in Black Howler Monkeys (Alouatta pigra) SARIE VAN BELLE1 AND ALEJANDRO ESTRADA

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