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		Downloaded from http://rspb.royalsocietypublishing.org/ on June 5, 2016 The Royal Society is collaborating with JSTOR to digitize, preserve, and extend access to Proceedings of the Royal Society of London. Series B, Bio Add to Reading List Source URL: rspb.royalsocietypublishing.org Download Document from Source Website File Size: 3,77 MB Share Document on Facebook